Due to the presence of a cell wall which surrounds and glues plant cells together, plants rely strongly on the interplay between oriented cell divisions and elongation to establish their overall body plan. Next to proliferative (symmetric) divisions, plants utilize formative (asymmetric) divisions to establish new organs. In the majority of higher plant cells, both proliferative and formative division planes are determined by the formation of a plant-specific cytoskeletal structure termed the preprophase band (PPB). This structure encircles the premitotic nucleus and determines the future division plane long before the cell actually divides. The PPB is however only a transient structure as it disappears prior to nuclear envelope breakdown and its exact function in establishing the future division plane remains still not fully understood.